#NEXUS

[ File saved by NDE version 0.4.7, Mon Oct 02 16:15:40 2000 ]

[!Project: Canarian_psyllids/]

BEGIN TAXA;
	DIMENSIONS NTAX=46;
	TAXLABELS
		'Acizzia uncatoides'
		'Arytinnis nigralineata'
		'Arytinnis pileolata'
		'Arytinnis hupalupa'
		'Arytinnis proboscidea'
		'Arytinnis romeria'
		'Arytinnis equitans'
		'Arytinnis prognata'
		'Arytinnis diluta'
		'Arytinnis occidentalis'
		'Arytinnis menceyata'
		'Arytinnis ochrita'
		'Arytinnis fortunata'
		'Arytinnis gomerae'
		'Arytinnis modica'
		'Arytinnis dividens'
		'Arytinnis hakani'
		'Arytinnis incuba'
		'Arytinnis cognata'
		'Arytinnis umbonata'
		'Arytinnis berber'
		'Arytaina devia'
		'Arytaina vittata'
		'Arytaina nubivaga'
		'Arytaina adenocarpi'
		'Arytaina genistae'
		'Arytainilla montivaga'
		'Arytainilla atlantica'
		'Arytainilla serpentina'
		'Arytainilla cytisi'
		'Arytainilla telonicola'
		'Arytainilla sulci'
		'Arytainilla spartiophila'
		'Arytainilla gredi'
		'Pseudacanthopsylla improvisa'
		'Livilla ima'
		'Livilla monospermae'
		'Livilla retamae'
		'Livilla spB'
		'Arytinnis canariensis'
		'Arytainilla delarbrei'
		'Livilla caprifuga'
		'Livilla baetica'
		'Livilla complexa'
		'Livilla spC'
		'Livilla nervosa'
		;
ENDBLOCK;

BEGIN CHARACTERS;
	DIMENSIONS NCHAR=67;
	FORMAT DATATYPE=STANDARD MISSING=? GAP=- SYMBOLS="0123456";
	CHARLABELS
		[1] 'wing shape'
		[2] 'costal break'
		[3] pterostigma
		[4] 'wing vein pigmentation'
		[5] 'termination of vein Rs at the wing apex'
		[6] 'superficial forewing spinules density'
		[7] 'shape of hindwing costal margin'
		[8] 'genal cones position'
		[9] 'genal cones terminal setae length'
		[10] 'genal cones terminal setae colour'
		[11] 'metatarsal spur'
		[12] 'male paramere shape'
		[13] 'male paramere sclerotized apex in lateral view'
		[14] 'male paramere sclerotized apex in dorsal view'
		[15] 'male paramere sclerotized apex inner surface in dorsal view'
		[16] 'male proctiger apex'
		[17] 'male adeagus distal segment apex'
		[18] 'tip of adeagus hook'
		[19] 'male subgenital plate dorsal profile'
		[20] 'male subgenital plate dorsal posterior margin'
		[21] 'female proctiger dorsal profile'
		[22] 'female proctiger apex'
		[23] 'female subgenital plate ventral profile'
		[24] 'female ovipositor'
		[25] 'female ovipositor valvulae dorsalis shape'
		[26] 'female ovipositor valvulae ventralis apex'
		[27] 'pterostigma length'
		[28] 'genal cone length'
		[29] 'antennal length'
		[30] 'distal proboscis segment length'
		[31] 'ratio forewing length to width'
		[32] 'ratio hind leg tibia length to femur length'
		[33] 'ratio hind leg apical tarsus to tibia length'
		[34] 'male proctiger length'
		[35] 'male paramere length'
		[36] 'male adeagus length'
		[37] 'male adeagus hook'
		[38] 'ratio male subgenital plate length to height'
		[39] 'ratio male paramere length to subgenital plate height'
		[40] 'female proctiger length'
		[41] 'female subgenital plate length'
		[42] 'female ventral ovipositor valve length'
		[43] 'ratio female proctiger length to circumanal ring length'
		[44] 'forewings and abdomen apically'
		[45] 'antennal segments'
		[46] 'circumanal ring shape'
		[47] 'body length'
		[48] 'antennae length'
		[49] 'ratio wing pad length to head width'
		[50] 'ratio caudal plate width to length'
		[51] 'ratio caudal plate width to circumanal ring width'
		[52] 'head setae'
		[53] 'antennal setae'
		[54] 'ocular seta'
		[55] 'secondary post-ocular seta'
		[56] 'dorsal thoracic setae'
		[57] 'dorsal prominent pre-caudal setae placement'
		[58] 'forewing pad setae'
		[59] 'forewing pad prominent setae'
		[60] 'forewing pad prominent setae'
		[61] 'abdominal sectasetae'
		[62] 'abdominal sectasetae'
		[63] 'marginal abdominal setae (other than sectasetae)'
		[64] 'leg setal type'
		[65] 'foreleg tibia distinct setal type'
		[66] 'middle leg tibia distinct setal type'
		[67] 'hind leg tibia distinct setal type'
		;
	STATELABELS
		1
			'widest in the apical third with a broadly rounded apex'
			'widest in the middle third with more acutely rounded apex'
			'margins more or less parallel, apex broadly rounded or more acute',
		2
			absent
			present,
		3
			absent
			present,
		4
			'uniformly coloured'
			'with distinct light and dark banding'
			'with dark patches at the terminal apices',
		5
			'strongly curved towards the wing margin'
			'not, or only weakly curved towards the wing margin',
		6
			'sparse: less than 40 per 0.1mm2'
			'medium density: 40-60 per 0.1mm2'
			'dense: 60-100 per 0.1mm2'
			'very dense: more than 100 per 0.1mm2',
		7
			straight
			'slightly concave'
			'markedly concave',
		8
			'commencing dorsally from more or less the same level as the vertex'
			'commencing ventrally relative to the vertex, with a distinct depression between the vertex and the genal cones',
		9
			'as long as, or typically longer than the vertex'
			'typically shorter than the vertex, occasionally as long',
		10
			'darkly pigmented'
			'not darkly pigmented',
		11
			absent
			one
			two,
		12
			'with an apically thin neck above a medially positioned blade or ridge produced on the external side and directed anteriorly'
			'apically thin neck typically absent or very short, without a medial blade but with a terminal ridge or blade present laterally exterior to the apex'
			'broad and parallel sided, lacking medial or apical blade'
			'slender, often simple with sides parrallel or tapering towards the apex, lacking medial or apical blade',
		13
			'rounded with small, or more pronounced, anteriorly, or interiorly directed hook'
			'flatened, or more or less so, with slight, or more pronounced, anterior projection'
			'either flatened, or with a well rounded bulge, and with slight anterior and posterior projections'
			'small, not hooked, but sometimes with small anterior projection; and displaced either to the interior or posterior of the top of the paramere',
		14
			'contiguous anteriorly'
			'contiguous more or less along entire inner margin'
			'contiguous posteriorly',
		15
			'straight edged'
			concave
			'rounded with an acute point',
		16
			'with pronounced posterior extension apically'
			'without pronounced posterior extension apically',
		17
			'with a well developed, curved hook'
			'with a somewhat flattened, shallow hook'
			'not developed into a hook',
		18
			'turning upwards'
			'not turning upwards',
		19
			'more or less straight and horizontal'
			'raised anteriorly, either with or without a distinct step',
		20
			'with a pair of long (more than half the length of the paramere) stout setae'
			'without a pair of long stout setae',
		21
			'more or less straight from anus to apex'
			'with a post anal depression, but subsequently more or less straight, or slightly concave or convex'
			'without a post anal depression, but concave with an upturned apex',
		22
			'squarely truncated'
			'bluntly rounded'
			acute,
		23
			'shallowly curved, without medial bulge, but sometimes angled medially'
			'slight, or more pronounced, medial bulge'
			'deeply curved, without medial bulge',
		24
			'valvulae ventralis slender, height of valvulae dorsalis typically, as great or, greater than valvulae ventralis'
			'valvulae ventralis robust, height of valvulae dorsalis less than valvulae ventralis',
		25
			'dorsally convex, wedge-shaped and tapering to the apex'
			'dorsally concave, and distinctly inflated anteriorly'
			'more or less straight, with or without dorsal, tooth-like processes'
			'triangular, tapering both to the apex and anteriorly',
		26
			'rounded, often acutely so, lacking any serration'
			'rounded, with minute serrations that extend along the ventral margin'
			'stepped with square tip'
			'rounded, with two small ventral processes near the tip',
		27
			'short, WLPT equal to or more than 20'
			'medium, WLPT 4.5-20'
			'long, WLPT equal to or less than 4.5',
		28
			'very long, more than 0.3 mm'
			'long, 0.15 - 0.3 mm'
			'short to very short, 0.15 mm or less',
		29
			'long, more than 2 mm'
			'medium, 1.5 - 2 mm'
			'short, 1 to 1.5 mm'
			'very short, less than 1 mm',
		30
			'up to 0.14 mm'
			'0.14 to 0.2 mm'
			'0.2 to 0.3 mm'
			'0.3 mm or more',
		31
			'greater than 2.6'
			'2.2-2.6'
			'less than 2.2',
		32
			'up to 1'
			'1 to 1.26'
			'1.26 or more',
		33
			'up to 0.22'
			'0.22 to 0.33'
			'0.33 or more',
		34
			'0.5 or greater'
			'0.4-0.5'
			'0.3-0.4'
			'0.3 or less',
		35
			'long, greater than 0.55 mm'
			'mid-length, greater than 0.4 but less than 0.55 mm'
			'short, 0.25 to 0.4 mm'
			'very short, less than 0.25 mm',
		36
			'up to 0.26'
			'0.26 to 0.37'
			'0.37 to 0.47'
			'0.47 or more',
		37
			'large, greater than 0.13 mm'
			'medium, 0.08-0.13 mm'
			'small, 0.08 mm or less',
		38
			'up to 1.3'
			'1.3 to 1.55'
			'1.55 or more',
		39
			'1.4 or greater'
			'1 to 1.4'
			'1 or less',
		40
			'1 mm or greater'
			'0.85 to 1 mm'
			'0.5 to 0.85 mm'
			'less than 0.5 mm',
		41
			'greater than 0.7 mm'
			'0.35 to 0.7 mm'
			'less than 0.35 mm',
		42
			'up to 0.2'
			'0.2 to 0.35'
			'0.35 or more',
		43
			'greater than 4'
			'4 or less',
		44
			acute
			'broadly rounded',
		45
			seven
			eight
			nine,
		46
			'broadly crescent-shaped with well rounded anterior lobes'
			'narrowly crescent-shaped with well rounded anterior lobes, but with lateral margins either straight or slightly concave'
			'narrowly crescent-shaped with acutely rounded anterior lobes',
		47
			'up to 2 mm'
			'2 mm or more',
		48
			'up to 0.63 mm'
			'0.63 to 0.88 mm'
			'0.88 to 1.18 mm'
			'1.18 mm or more',
		49
			'greater than 1'
			'1 or less',
		50
			'greater than 1.2'
			'less than 1.2',
		51
			'up to 4'
			'4 to 5'
			'5 or more',
		52
			'simple, occasionally narrowly capitate'
			'simple and distinctly capitate'
			'simple and club-shaped'
			'simple and sectasetae',
		53
			simple
			'simple and capitate'
			'simple and club-shaped'
			sectasetae,
		54
			absent
			'simple, or narrowly capitate or rod'
			'distinctly capitate'
			sectasetae,
		55
			'absent or indistinct'
			'prominent, simple, sometimes narrowly capitate'
			'prominent, distinctly capitate, occasionally narrow',
		56
			'simple only'
			'capitate present'
			'club setae'
			sectasetae,
		57
			'absent or indistinct'
			'prominent, on anterior tergites'
			'prominent, on posterior tergites'
			'prominent, on all tergites',
		58
			'simple only'
			'capitate only'
			'simple and capitate'
			'club-shaped'
			sectasetae,
		59
			marginal
			'surface and marginal',
		60
			one
			two
			'more than two',
		61
			absent
			present,
		62
			'one pair'
			'two pairs'
			'three pairs'
			'four pairs'
			'more than four pairs',
		63
			absent
			'one pair'
			'two pairs'
			'three or four pairs'
			'more than four pairs',
		64
			'simple setae only'
			'capitate setae present'
			'club-shaped setae present'
			'sectasetae present',
		65
			one
			two
			'more than two',
		66
			one
			two
			'more than two',
		67
			one
			two
			'more than two',
	;
	MATRIX
		'Acizzia uncatoides'              0110121011 1322012101 2100032230 1123301213 2011200010 2112013112 0-31222
		'Arytinnis nigralineata'          0111101100 1111011011 01000322{01}{23} 12011{12}1010 0101200{23}10 0001111201 1230---
		'Arytinnis pileolata'             0110101101 1022011101 01100022{12}1 1{12}13211{01}12 1011000210 0001211102 1331-01
		'Arytinnis hupalupa'              0110101101 1030010011 0100002220 1112011101 1101000110 {01}00110{01}{02}00 1331-0{01}
		'Arytinnis proboscidea'           0111101100 1021011111 0100032213 12021{12}1010 0101200210 {01}001101001 1230---
		'Arytinnis romeria'               0110111101 1300211101 0100002221 1113202{01}12 1011000110 {01}001201102 1331-00
		'Arytinnis equitans'              0110111101 1132011101 0110002212 12{01}3211012 1011000210 0001{12}11201 123{01}--0
		'Arytinnis prognata'              0110111101 1311011101 2110002221 1{12}1{23}211{01}12 100100{01}{12}10 0001101200 1310---
		'Arytinnis diluta'                0110121100 1130011001 110000222{01} 1{12}{01}2112{01}02 1001000110 1001102000 1330---
		'Arytinnis occidentalis'          0110101101 1300110111 21000022{01}1 1113211012 1011100210 0001101201 1330---
		'Arytinnis menceyata'             0110121101 1022110111 0100002221 1112011101 1101000110 1001113212 1331012
		'Arytinnis ochrita'               0110111101 1011011101 0100002220 1113202102 10{01}1000110 000?000100 1231--0
		'Arytinnis fortunata'             0110111101 1301011101 0100002211 1113202012 1011000210 0001101202 1331-02
		'Arytinnis gomerae'               0110101101 1310111111 2100002220 1113212012 1011100{12}10 000121120{12} 1{23}30---
		'Arytinnis modica'                0110101101 1030010011 0100002211 11110{12}1100 0101000{12}10 {01}001101200 1331-1{12}
		'Arytinnis dividens'              0110101101 1030010011 01000022{01}{12} 1111021{01}00 010100{01}{23}10 {012}001{12}11201 1{23}31-{12}2
		'Arytinnis hakani'                0110101101 1012011111 0100002211 1{12}131{01}1012 10{01}1000210 0001{12}11202 1230---
		'Arytinnis incuba'                0110101101 1012011111 0100002211 1112111011 1001000{23}10 0001211202 1230---
		'Arytinnis cognata'               0110101101 1022011111 0110002211 1113111012 1011000210 {01}001101202 1330---
		'Arytinnis umbonata'              0110101101 1130211111 2110002211 111{23}2{01}2{01}22 1011000210 {01}001211202 1230---
		'Arytinnis berber'                0110101101 1010011101 0100002221 1113202{01}12 1011000110 0001211202 1330---
		'Arytaina devia'                  0002101101 1100201101 110000-21{12} 11122120{12}2 1011000210 {01}001211112 {01}031122
		'Arytaina vittata'                20001021{01}1 1130100101 110000-100 0021211{01}{12}{01} 111100{01}{23}10 0001101200 1031-00
		'Arytaina nubivaga'               20001021{01}1 1130100101 110000-101 002{01}211021 1111000{23}10 100110121{01} 1131-22
		'Arytaina adenocarpi'             1002101101 1200100101 110000-2{12}1 111{12}2{01}2{01}1{12} 1{01}1100{01}{12}10 {01}0112{01}3212 1131122
		'Arytaina genistae'               1002122101 1200100101 110000-111 0022211011 111100{01}210 1011{12}03212 {01}{01}31{12}22
		'Arytainilla montivaga'           0110100101 1311011101 {01}100001221 1112{12}{01}1012 101100{01}110 1111213212 0-31122
		'Arytainilla atlantica'           1110110101 1300011101 0101221220 111{12}11{12}002 12010200{01}0 {12}011213012 1331{12}22
		'Arytainilla serpentina'          1110120011 1300212101 010120{12}120 0120032000 0200001101 10010000-- 1220---
		'Arytainilla cytisi'              1110110111 1300011101 0101221220 1111022000 020002{01}100 {12}221023300 1332222
		'Arytainilla telonicola'          0110110111 1300011101 0101221220 1112112{01}01 12000201{01}1 {12}221023300 1332222
		'Arytainilla sulci'               0110120011 1100211111 102111{12}120 1121121012 1111011{12}10 2010213112 1041222
		'Arytainilla spartiophila'        0110130111 1300211101 2221301220 1{12}121120{01}2 12{01}10000{01}0 0101000100 1331-22
		'Arytainilla gredi'               0110110001 1300212101 2210000220 1112212112 1111000{01}{01}0 1001103202 1331-{01}{01}
		'Pseudacanthopsylla improvisa'    2110111011 0300011101 1200002120 1023202012 1010020010 2333033412 1403222
		'Livilla ima'                     0110110011 1100010101 0100001110 1111120{01}10 11{01}1000{12}00 1001{01}00200 1331-00
		'Livilla monospermae'             2110030011 0300211100 1100000000 0111121010 011100{01}210 0112211112 1331222
		'Livilla retamae'                 2110030011 0300211100 210000{01}100 0111121010 011100{01}{23}10 {01}111{12}111{01}2 1331{012}22
		'Livilla spB'                     2110030011 1300211101 1100000110 1112202{01}12 10110001{01}0 0001111202 1331-{01}{01}
		'Arytinnis canariensis'           0110101100 1022011101 0100002212 1202211012 101??????? ?????????? ???????
		'Arytainilla delarbrei'           11101{12}0011 1300?11101 2211201111 1110032000 020??????? ?????????? ???????
		'Livilla caprifuga'               1000130011 1300211111 010000-121 1112111011 111??????? ?????????? ???????
		'Livilla baetica'                 0110130001 1311011111 0100001121 1111120010 010??????? ?????????? ???????
		'Livilla complexa'                1010130011 1300211101 110000112{01} 2112211012 101??????? ?????????? ???????
		'Livilla spC'                     2110030011 1300211101 2100000110 {01}212212012 111??????? ?????????? ???????
		'Livilla nervosa'                 1000030011 1300211101 010000-110 02112{12}1110 010??????? ?????????? ???????
	;
ENDBLOCK;

BEGIN ASSUMPTIONS;
	OPTIONS DEFTYPE=UNORD POLYTCOUNT=MINSTEPS;
	TYPESET * default = ORD: 6 27-42 47-51;
ENDBLOCK;

BEGIN NOTES;
	[Taxon comments]

	[Character comments]

	[Character state comments]
	TEXT CHARACTER=1 STATE=0 TEXT='lacking distinct surface pattern, not coriaceous';
	TEXT CHARACTER=1 STATE=1 TEXT='lacking distinct surface pattern, but if coriaceous more darkly pigmented towards the margins ';
	TEXT CHARACTER=1 STATE=2 TEXT='with or without distinct surface pattern, not coriaceous';
	TEXT CHARACTER=27 STATE=0 TEXT='less than one 20th the wing length';
	TEXT CHARACTER=27 STATE=1 TEXT='less than one quarter of the wing length';
	TEXT CHARACTER=27 STATE=2 TEXT='about one quarter of the wing length';
	TEXT CHARACTER=28 STATE=0 TEXT='about 1.5 times the length of the vertex';
	TEXT CHARACTER=28 STATE=1 TEXT='roughly the same length as the vertex';
	TEXT CHARACTER=28 STATE=2 TEXT='up to half the length of the vertex';

	[Attribute comments]

	[Taxon pictures]

	[Character pictures]

	[Character state pictures]

	[Attribute pictures]
ENDBLOCK;